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Israeli troops opened fire Saturday toward crowds of Palestinians seeking food from distribution hubs run by a  in southern Gaza, killing 32 people, according to witnesses and hospital officials.

The two incidents occurred near hubs operated by the Gaza Humanitarian Foundation. In other violence, two Palestinians were killed in an Israeli airstrike in Gaza City, health officials said.

The GHF  with backing from the U.S. and Israel. The two governments are seeking to replace the traditional U.N.-led aid distribution system in Gaza, saying that Hamas militants siphon off supplies. The U.N. denies the allegation.

While the GHF says it has distributed millions of meals to hungry Palestinians, local health officials and witnesses say that hundreds of people have been  as they try to reach the distribution hubs.

The army, which is not at the sites but secures them from a distance, says it only fires warning shots if crowds get too close to its forces.

The GHF, which employs private armed guards, says there have been no deadly shootings at its sites, though this week, 20 people were killed at one of its locations, most of them in a stampede. The group accused Hamas agitators of causing a panic, but gave no evidence to back the claim.

In a statement, the GHF said there were no incidents at or near its sites. It said the reported Israeli shootings occurred far from the sites and hours before they opened. "We have repeatedly warned aid seekers not to travel to our sites overnight and early morning hours," it said.

The Israeli military said it had fired "warning shots" near Rafah after a group of suspects approached troops and ignored calls to keep their distance. It said it was investigating reports of casualties, but noted the incident occurred overnight when the distribution site was closed.

Most of Saturday's deaths occurred as Palestinians massed in the Teina area, around two miles away from a GHF aid distribution center east of the city of Khan Younis.

Mahmoud Mokeimar, an eyewitness, said he was walking with masses of people — mostly young men — toward the food hub. Troops fired warning shots as the crowds advanced, before opening fire toward the marching people.

"It was a massacre … the occupation opened fire at us indiscriminately," he said. He said he managed to flee but saw at least three motionless bodies lying on the ground, and many other wounded fleeing.

Akram Aker, another witness, said troops fired machine guns mounted on tanks and drones. He said the shooting happened between 5 a.m. and 6 a.m.

"They encircled us and started firing directly at us," he said. He said he saw many casualties lying on the ground.

Sanaa al-Jaberi, a 55-year-old woman, said she saw many dead and wounded as she fled the area.

"We shouted: 'food, food,' but they didn't talk to us. They just opened fire," she said.

Monzer Fesifes, a Palestinian-Jordanian, said his 19-year-old son Hisham was among those killed in the Teina area.

"He went to bring food from the failed US, Zionist aid to feed us," the father of six said, pleading for the Jordanian government to help evacuate them from the Palestinian enclave.

The Nasser Hospital in Khan Younis said it received 25 bodies, along with dozens wounded.

Seven other people, including one woman, were killed in the Shakoush area, hundreds of meters (yards) north of another GHF hub in Gaza's southernmost city of Rafah, the hospital said. The toll was also confirmed by the health ministry.

Dr. Mohamed Saker, the head of Nasser's nursing department, said it received 70 wounded people. He told The Associated Press that most of the casualties were shot in their heads and chests, and that some were placed in the already overwhelmed intensive care unit.

"The situation is difficult and tragic," he said, adding that the facility lacks badly needed medical supplies to treat the daily flow of casualties.

Meanwhile, Fares Awad, head of the Health Ministry's ambulance and emergency service in northern Gaza, confirmed the two deaths in Gaza City. He said an airstrike hit a tent in a camp sheltering displaced families in the courtyard of the Development Ministry. There were no further details on the target of the strike.

The Israeli army said it had struck some 90 targets throughout Gaza over the past day and that it had killed militants and targeted "terror infrastructure" in northern Gaza and Gaza City.

Gaza's more than 2 million Palestinians are living through a catastrophic humanitarian crisis, and the territory is teetering on the edge of famine, according to food security experts.

Distribution at the GHF sites has often been chaotic. Boxes of food are left stacked on the ground inside the centers and, once opened, crowds charge in to grab whatever they can, according to witnesses and videos released by GHF itself.

In videos obtained recently by the AP from an American contractor working with GHF, contractors are seen using tear gas and stun grenades to keep crowds behind metal fences or to force them to disperse. Gunshots can also be heard.

Hamas triggered the ongoing 21-month war in Gaza when terrorists on Oct. 7, 2023, killing some 1,200 people and taking .

An Israeli military offensive has killed over 58,000 Palestinians, according to the Hamas-run Gaza Health Ministry, displaced nearly all of Gaza's 2 million people and caused widespread destruction.

The ministry does not say how many militants are among the dead, though it says over half were women and children. The ministry is part of the Hamas government, but is seen by the U.N. and other international organizations as the most reliable source of data on war casualties.

Israel and Hamas have been holding in recent weeks. But international mediators say there have been no breakthroughs in the talks.